The main objective of Quake IV, which is revealed over the course of the game, is the destruction of the Nexus, a facility believed by Human commanders to be essential for communication between all Strogg units, giving them a tremendous advantage over invaders. After the Strogg planetary defense systems have been disabled and the Makron has been killed by a lone survivor of the first invasion, humanity now leads a massive onslaught on the Strogg homeworld. Quake IV takes place during the Second Invasion of Stroggos. In fact, your worst nightmare is soon realized when you discover to defeat the Strogg, you must become one of them. Equipped with hi-tech weapons and mobile vehicles, and teamed with a top Marine squadron, you soon realize this war is far from over. You are Matthew Kane, a member of the elite Rhino Squad and part of the next massive invasion that spans across the entire Stroggos planet. The Strogg proved to be extremely resilient and as the war continued to rage, the Strogg regrouped and constructed a new and more powerful Makron. However, one surviving Marine broke through and was able to destroy The Big Gun and ultimately defeat the Strogg leader, the Makron. The first assault force was launched from the fleet and encountered massive resistance - most of the drop-ships were destroyed by the Strogg homeworld's massive defense weapon known as 'The Big Gun'. The Strogg battled the human race for years on Earth, but as resources became depleted, Earth mounted a counter-assault on the Strogg homeworld. Through a horrific process, the limbs and flesh of the fallen are fused with metal and machinery to create the monstrosities that are the alien Strogg. While the invasion was originally believed to be an attempt to strip the planet of its natural resources, humans soon discovered the Strogg are only after one thing: the human remains of those they kill. In the mid-21st century, a barbaric alien race called the Strogg launched a massive invasion on Earth.

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